The giant panda ( Ailuropoda melanoleuca), also known as the panda bear or simply panda, is a bear species endemic to China. It is characterised by its white animal coat with black patches around the eyes, ears, legs and shoulders. Its body is rotund; adult individuals weigh and are typically long. It is sexually dimorphic, with males being typically 10 to 20% larger than females. A thumb is visible on its forepaw, which helps in holding bamboo in place for feeding. It has large molar teeth and expanded temporal fossa to meet its dietary requirements. It can digest starch and is mostly herbivorous with a diet consisting almost entirely of bamboo and .
The giant panda lives exclusively in six montane regions in a few Chinese provinces at elevations of up to . It is solitary and gathers only in mating seasons. It relies on olfactory communication to communicate and uses as chemical cues and on landmarks like rocks or trees. Females rear cubs for an average of 18 to 24 months. The oldest known giant panda was 38 years old.
As a result of farming, deforestation and infrastructural development, the giant panda has been driven out of the lowland areas where it once lived. The Fourth National Survey (2011–2014), published in 2015, estimated that the wild population of giant pandas aged over 1.5 years (i.e. excluding dependent young) has increased to 1,864 individuals, based on this number and using the available estimated percentage of cubs in the population (9.6%) the IUCN estimated the total number of Pandas to be approximately 2,060. Since 2016, it has been listed as Vulnerable on the IUCN Red List. In July 2021, Chinese authorities also classified the giant panda as vulnerable. It is a conservation-reliant species. By 2007, the captive population comprised 239 giant pandas in China and another 27 outside the country. It has often served as China's national symbol, appeared on Chinese Gold Panda coins since 1982 and as one of the five Fuwa mascots of the 2008 Summer Olympics held in Beijing.
Since the earliest collection of Chinese writings, the Chinese language has given the bear many different names, including mò (貘, ancient Chinese name for giant panda),Harper, Donald (2013), "The Cultural History of the Giant Panda (Ailuropoda melanoleuca) in Early China", Early China 35/36: 185–224. ( abstract ) huāxióng (花熊; "spotted bear") and zhúxióng (竹熊; "bamboo bear"). The most popular names in China today are dàxióngmāo (大熊貓; ), or simply xióngmāo (熊貓; ). As with the word panda in English, xióngmāo (熊貓) was originally used to describe just the red panda, but dàxióngmāo (大熊貓) and xiǎoxióngmāo (小熊猫; ) were coined to differentiate between the species.
In Taiwan, another popular name for panda is the inverted dàmāoxióng (大貓熊; ), though many encyclopedias and dictionaries in Taiwan still use the "bear cat" form as the correct name. Some linguists argue, in this construction, "bear" instead of "cat" is the base noun, making the name more grammatically and logically correct, which have led to the popular choice despite official writings. This name did not gain its popularity until 1988, when a private zoo in Tainan painted a sun bear black and white and created the Tainan fake panda incident.
A detailed study of the giant panda's genetic history from 2012 confirms that the separation of the Qinling population occurred about 300,000 years ago, and reveals that the non-Qinling population further diverged into two groups, named the Minshan and the Qionglai-Daxiangling-Xiaoxiangling-Liangshan group respectively, about 2,800 years ago.
The species has been located at elevations of above sea level. They frequent habitats with a healthy concentration of bamboos, typically old-growth forests, but may also venture into secondary forest habitats. The Daxiangling Mountain population inhabits both coniferous and broadleaf forests. Additionally, the Qinling population often selects evergreen broadleaf and conifer forests, while pandas in the Qionglai mountainous region exclusively select upland conifer forests. The remaining two populations, namely those occurring in the Liangshan and Xiaoxiangling mountains, predominantly occur in broadleaf evergreen and conifer forests.
Giant pandas once roamed across Southeast Asia from Myanmar to northern Vietnam. Their range in China spanned much of the southeast region. By the Pleistocene, climate change affected panda populations, and the subsequent domination of led to large-scale habitat loss. In 2001, it was estimated that the range of the giant panda had declined by about 99% of its range in earlier millenniums.
The panda's skull shape is typical of durophagous carnivorans. It has evolved from previous ancestors to exhibit larger molars with increased complexity and expanded temporal fossa. A study revealed that a giant panda had a bite force of 1298.9 Newton (BFQ 151.4) at canine teeth and 1815.9 Newton (BFQ 141.8) at carnassial teeth.
Adults measure around long, including a tail of about , and tall at the shoulder. Males can weigh up to . Females are generally 10–20% smaller than males. They weigh between and . The average weight for adults is .
The giant panda's paw has a digit similar to a thumb and five fingers; the thumb-like digit – actually a modified sesamoid bone – helps it to hold bamboo while eating. The giant panda's tail, measuring , is the second-longest in the bear family, behind the sloth bear.
During the shoot season (AprilAugust), pandas store a large amount of food in preparation for the months succeeding this seasonal period, in which pandas live off a diet of bamboo leaves. The giant panda is a highly specialised animal with unique adaptations, and has lived in bamboo forests for millions of years.
The average giant panda eats as much as of bamboo shoots a day to compensate for the limited energy content of its diet. Ingestion of such a large quantity of material is possible and necessary because of the rapid passage of large amounts of indigestible plant material through the short, straight digestive tract. It is also noted, however, that such rapid passage of digesta limits the potential of microbial digestion in the gastrointestinal tract, limiting alternative forms of digestion. Given this voluminous diet, the giant panda defecates up to 40 times a day. The limited energy input imposed on it by its diet has affected the panda's behavior. The giant panda tends to limit its social interactions and avoids steeply sloping terrain to limit its energy expenditures.
Two of the panda's most distinctive features, its large size and round face, are to its bamboo diet. Anthropologist Russell Ciochon observed: "much like the vegetarian gorilla, the low body surface area to body volume of is indicative of a lower metabolic rate. This lower metabolic rate and a more sedentary lifestyle allows the giant panda to subsist on nutrient poor resources such as bamboo." The giant panda's round face is the result of powerful jaw muscles, which attach from the top of the head to the jaw. Large molars crush and grind fibrous plant material.
The morphological characteristics of extinct relatives of the giant panda suggest that while the ancient giant panda was omnivorous 7 million years ago (mya), it only became herbivorous some 2–2.4 mya with the emergence of A. microta. Genome sequencing of the giant panda suggests that the dietary switch could have initiated from the loss of the sole umami taste receptor, encoded by the genes TAS1R1 and TAS1R3 (also known as T1R1 and T1R3), resulting from two frameshift mutations within the T1R1 exons. Umami taste corresponds to high levels of glutamate as found in meat and may have thus altered the food choice of the giant panda. Although the pseudogenisation (conversion into a pseudogene) of the umami taste receptor in Ailuropoda coincides with the dietary switch to herbivory, it is likely a result of, and not the reason for, the dietary change. The mutation time for the T1R1 gene in the giant panda is estimated to 4.2 mya while fossil evidence indicates bamboo consumption in the giant panda species at least 7 mya, signifying that although complete herbivory occurred around 2 mya, the dietary switch was initiated prior to T1R1 loss-of-function.
Pandas eat any of 25 bamboo species in the wild, with the most common including Fargesia dracocephala and Fargesia rufa. Only a few bamboo species are widespread at the high altitudes pandas now inhabit. Bamboo leaves contain the highest protein levels; stems have less. Because of the synchronous flowering, death, and regeneration of all bamboo within a species, the giant panda must have at least two different species available in its range to avoid starvation. While primarily herbivorous, the giant panda still retains decidedly ursine teeth and will eat meat, fish, and eggs when available. In captivity, zoos typically maintain the giant panda's bamboo diet, though some will provide specially formulated biscuits or other dietary supplements. (page numbers as per the 2002 edition)
Pandas will travel between different habitats if they need to, so they can get the nutrients that they need and to balance their diet for reproduction.
Giant pandas are Sympatry with other large mammals and bamboo feeders, such as the takin ( Budorcas taxicolor). The takin and giant panda share a similar ecological niche, and they consume the same resources. When competition for food is fierce, pandas disperse to the outskirts of takin distribution. Other possible competitors include but is not limited to, the Wild boar ( Sus scrofa), Chinese goral ( Naemorhedus griseus) and the Asian black bear ( Ursus thibetanus). Giant pandas avoid areas with a mid-to-high density of livestock, as they depress the vegetation. The Tibetan Plateau is the only known area where both giant and can be found. Although sharing near-identical ecological niches, competition between the two species has rarely been observed. Nearly 50% of their respective distribution overlaps, and successful coexistence is achieved through distinct habitat selection.
Pandas communicate through vocalisation and scent marking such as clawing trees or spraying urine. They are able to climb and take shelter in hollow trees or rock crevices, but do not establish permanent dens. For this reason, pandas do not hibernation, which is similar to other subtropical mammals, and will instead move to elevations with warmer temperatures. Pandas rely primarily on spatial memory rather than visual memory. Though the panda is often assumed to be docile, it has been known to attack humans on rare occasions. Pandas have been known to cover themselves in horse manure to protect themselves against cold temperatures.
The species communicates foremost through a blatting sound; they achieve peaceful interactions through the emission of this sound. When in oestrus, a female emits a chirp. In hostile confrontations or during fights, the giant panda emits vocalizations such as a roar or growl. On the other hand, squeals typically indicate inferiority and submission in a dispute. Other vocalizations include honks and moans.
The season plays a major role in mediating chemical communication. Depending on the season, mainly whether it is breeding season or not, may influence which odors are prioritized. Chemical signals can have different functions in different seasons. During the non-breeding season, females prefer the odors of other females because reproduction is not their primary motivation. However, during breeding season, odors from the opposite sex will be more attractive. Because they are solitary mammals and their breeding season is so brief, female pandas secrete chemical cues in order to let males know their sexual status. The chemical cues female pandas secrete can be considered to be for sexual reproduction. Females deposit scent marks through their urine which induces an increase in androgen levels in males. Androgen is a sex hormone found in both males and females; testosterone is the major androgen produced by males. Civetone and decanoic acid are chemicals found in female urine which promote behavioral responses in males; both chemicals are considered giant panda pheromones. Male pandas also secrete chemical signals that include information about their sexual reproductivity and age, which is beneficial for a female when choosing a mate. For example, age can be useful for a female to determine sexual maturity and sperm quality.
Chemical cues, or odors, play an important role in how a panda chooses their habitat. Pandas look for odors that tell them not only the identity of another panda, but if they should avoid them or not. Pandas tend to avoid their species for most of the year, breeding season being the brief time of major interaction. Chemical signaling allows for avoidance and competition. Pandas whose habitats are in similar locations will collectively leave scent marks in a unique location which is termed "scent stations". When pandas come across these scent stations, they are able to identify a specific panda and the scope of their habitat. This allows pandas to be able to pursue a potential mate or avoid a potential competitor.
Pandas can assess an individual's dominance status, including their age and size, via odor cues and may choose to avoid a scent mark if the signaler's competitive ability outweighs their own. A panda's size can be conveyed through the height of the scent mark. Since larger animals can place higher scent marks, an elevated scent mark advertises a higher competitive ability. Age must also be taken into consideration when assessing a competitor's fighting ability. For example, a mature panda will be larger than a younger, immature panda and possess an advantage during a fight.
When the cub is first born, it is pink, blind, and toothless, weighing only , or about of the mother's weight, proportionally the smallest baby of any placental mammal. Guinness World Records 2013, Page 050, hardcover edition. It nurses from its mother's breast six to 14 times a day for up to 30 minutes at a time. For three to four hours, the mother may leave the den to feed, which leaves the cub defenseless. One to two weeks after birth, the cub's skin turns grey where its hair will eventually become black. Slight pink colour may appear on the cub's fur, as a result of a chemical reaction between the fur and its mother's saliva. A month after birth, the colour pattern of the cub's fur is fully developed. Its fur is very soft and coarsens with age. The cub begins to crawl at 75 to 80 days; mothers play with their cubs by rolling and wrestling with them. The cubs can eat small quantities of bamboo after six months, though mother's milk remains the primary food source for most of the first year. Giant panda cubs weigh at one year and live with their mothers until they are 18 months to two years old. The interval between births in the wild is generally two years.
Initially, the primary method of breeding giant pandas in captivity was by artificial insemination, as they seemed to lose their interest in mating once they were captured. This led some scientists to trying methods such as showing them videos of giant pandas mating and giving the males sildenafil (commonly known as Viagra). In the 2000s, researchers started having success with captive breeding programs, and they have now determined giant pandas have comparable breeding to some populations of the American black bear, a thriving bear species.
In July 2009, Chinese scientists confirmed the birth of the first cub to be successfully conceived through artificial insemination using frozen sperm. The technique for freezing the sperm in liquid nitrogen was first developed in 1980 and the first birth was hailed as a solution to the dwindling availability of giant panda semen, which had led to inbreeding. Panda semen, which can be frozen for decades, could be shared between different zoos to save the species. As of 2009, it is expected that zoos in destinations such as San Diego in the United States and Mexico City will be able to provide their own semen to inseminate more giant pandas.
Attempts have also been made to reproduce giant pandas by interspecific pregnancy where cloned panda embryos were implanted into the uterus of an animal of another species. This has resulted in panda fetuses, but no live births.
The creature named mo (貘) mentioned in some ancient books has been interpreted as giant panda. The dictionary Shuowen Jiezi (Eastern Han Dynasty) says that the mo, from Shu (Sichuan), is bear-like, but yellow-and-black, Shuowen Jiezi, Chapter 10, radical 豸: "貘:似熊而黃黑色,出蜀中" (" Mo: like bear, but yellow-and-black, comes from Shu"). although the older Erya describes mo simply as a "white leopard". Erya, Chapter "釋獸" ("About animals") : "貘,白豹" ( Mo, white leopard). The interpretation of the legendary fierce creature pixiu (貔貅) as referring to the giant panda is also common. China Giant Panda Museum: Historical Records in Ancient China . Supposed Chinese historical terminology appears in the Chinese version of this article, 我国古代的历史记载
During the reign of the Yongle Emperor (early 15th century), his relative from Kaifeng sent him a captured zouyu (騶虞), and another zouyu was sighted in Shandong. Zouyu is a legendary "righteous" animal, which, similarly to a qilin, only appears during the rule of a benevolent and sincere monarch.
the 1970s, gifts of giant pandas to American and Japanese zoos formed an important part of the diplomacy of the People's Republic of China (PRC), as it marked some of the first cultural exchanges between China and the West. This practice has been termed "panda diplomacy". By 1984, however, pandas were no longer given as gifts. Instead, China began to offer pandas to other nations only on 10-year loans for a fee of up to US$1,000,000 per year and with the provision that any cubs born during the loan are the property of China. As a result of this change in policy, nearly all the pandas in the world are owned by China, and pandas leased to foreign zoos and all cubs are eventually returned to China. As of 2022, Xin Xin at the Chapultepec Zoo in Mexico City, was the last living descendant of the gifted pandas.
Since 1998, because of a WWF lawsuit, the United States Fish and Wildlife Service only allows US zoos to import a panda if the zoo can ensure China channels more than half of its loan fee into conservation efforts for giant pandas and their habitat. In May 2005, China offered a breeding pair to Taiwan. The issue became embroiled in cross-Strait relations – due to both the underlying symbolism and technical issues such as whether the transfer would be considered "domestic" or "international" or whether any true conservation purpose would be served by the exchange. China's Panda Politics. Newsweek. 15 October 2007. Retrieved 23 May 2008. A contest in 2006 to name the pandas was held in the mainland, resulting in the politically charged names Tuan Tuan and Yuan Yuan (from l=reunion, implying reunification). China's offer was initially rejected by Chen Shui-bian, then President of Taiwan. However, when Ma Ying-jeou assumed the presidency in 2008, the offer was accepted and the pandas arrived in December of that year. China sends panda peace offering . The Guardian. 28 December 2008.
In the 2020s, certain "celebrity pandas" have gained a cult following amongst internet users, with dedicated fan accounts existing to keep tabs on the animals. Known as "giant panda fever" or "panda-monium", individual pandas are known to get billions of views and engagements on social media, as well as product lines specifically emulating them. At Chengdu Research Base of Giant Panda Breeding, certain of these "celebrity pandas" are known to garner hours-long lines specifically to see them.
The giant panda has been a target of poaching by locals since ancient times and by foreigners since it was introduced to the West. Starting in the 1930s, foreigners were unable to poach giant pandas in China because of the Second Sino-Japanese War and the Chinese Civil War, but pandas remained a source of soft furs for the locals. The population boom in China after 1949 created stress on the pandas' habitat and the subsequent famines led to the increased hunting of wildlife, including pandas. After the Chinese economic reform, demand for panda skins from Hong Kong and Japan led to illegal poaching for the black market, acts generally ignored by the local officials at the time. In 1963, the PRC government set up Wolong National Nature Reserve to save the declining panda population.
The giant panda is among the world's most adored and protected rare animals, and is one of the few in the world whose natural inhabitant status was able to gain a UNESCO World Heritage Site designation. The Sichuan Giant Panda Sanctuaries, located in the southwest province of Sichuan and covering seven natural reserves, were inscribed onto the World Heritage List in 2006. A 2015 paper found that the giant panda can serve as an umbrella species as the preservation of their habitat also helps other endemic species in China, including 70% of the country's forest birds, 70% of mammals and 31% of amphibians.
In 2012, Earthwatch Institute, a global nonprofit that teams volunteers with scientists to conduct important environmental research, launched a program called "On the Trail of Giant Panda". This program, based in the Wolong National Nature Reserve, allows volunteers to work up close with pandas cared for in captivity, and help them adapt to life in the wild, so that they may breed, and live longer and healthier lives. Efforts to preserve the panda bear populations in China have come at the expense of other animals in the region, including snow leopards, wolves, and dholes. In order to improve living and mating conditions for the fragmented populations of pandas, nearly 70 natural reserves have been combined to form the Giant Panda National Park in 2020. With a size of 10,500 square miles, the park is roughly three times as large as Yellowstone National Park and incorporates the Wolong National Nature Reserve. Small, isolated populations run the risk of inbreeding and smaller genetic variety makes the individuals more vulnerable to various defects and genetic mutation.
In 2020, the panda population of the new national park was already above 1,800 individuals, which is roughly 80 percent of the entire panda population in China. Establishing the new protected area in the Sichuan also gives various other endangered or threatened species, like the Siberian tiger, the possibility to improve their living conditions by offering them a habitat. Other species who benefit from the protection of their habitat include the snow leopard, the golden snub-nosed monkey, the red panda and the complex-toothed flying squirrel.
In July 2021, Chinese conservation authorities announced that giant pandas are no longer endangered in the wild following years of conservation efforts, with a population in the wild exceeding 1,800. China has received international praise for its conservation of the species, which has also helped the country establish itself as a leader in endangered species conservation.
Ecology
Diet
Interspecific interactions
Pathogens and parasites
Behavior
Olfactory communication
Reproduction
Human interaction
Early references
In captivity
Conservation
Population
See also
External links
|
|